Dan is a mathematician, engineer and physician. He completed his DPhi in mathematics at Oxford under the supervision of Prof Philip Maini FRS and his medical degree at Green-Templeton College, Oxford. He was a Miller Postdoctoral Fellow at UC Berkeley in the laboratory of MAR Koehl. Dan's research focusses on using mathematics and computer science methods to better understand complex biological systems, particularly as they relate to disease states, inflammation and ageing. A unifying theme of this work is viewing immune networks as devices of computation and control, with disease states disturbing the native topology.